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Crown of a Psaronius tree

Reconstruction of the leafy crown of a Psaronius tree by Stidd (1971).

From research on coal balls from the USA it is known that the leaves of Psaronius are of the Pecopteris-type because petioles with attached leaves has been found. Many of these leaflets still have the sporangia. These are fused in a characteristic way into synangia. Leaves of Pecopteris bearing such synangia are called Scolecopteris (at the bottom of the drawing).