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Two floras

Two groups of fossil floras can be distinguished:
I. floras representing marsh vegetations
II. floras representing vegetations growing outside the actual sedimentation basin
As to the second group plant remains deposited via a river in a lake or delta come to mind.
The known floras of the Upper Carboniferous belong nearly all to the first group, those of the Permian mainly to the second one. The possibilities for fossilisation in the first group were much better than those in the second group. The beautifully preserved and abundant coal fossils prove this.
In the second group the plant remains covered a long distance in the water before they were deposited in the sediment. That is the reason they are often very fragmentary.

The sediment with the Lodève flora is a lake deposition. Especially the tough plant remains, like branches of conifers are well represented. Ferns are very rare: these could obviously not stand the transportation. Small conifer twigs could have been blown into the water during storms.
The floras of the Carboniferous generally grew in a very humid environment. The conifer dominated Permian floras represent a completely different environment. Generally speaking: the number of humidity loving vegetations decreased strongly in the course of time and the vegetations adapted to dryer circumstances came more and more to the forefront in the Permian.